Contributing Authors
Contributing Authors
Authorship of accounts is by invitation only, but we encourage any and all who have expertise on particular turtle or tortoise species or subspecies to contact us about volunteering to take on (as senior author) or join (as co-author) a new account or an account in preparation for which you may have information to provide, or eventually, assisting or leading on a revised account. See specific instructions for Contributing Authors. Thank you.
To date, the following individuals have contributed published accounts in this on-going project
Number of contributing authors published to date: 250
Number of accounts published to date: 120 plus 9 checklists
Godfrey C. Akani
• Pelusios niger, 105
Erika A. Alacs
• Carettochelys insculpta, 009
• Chelodina burrungandjii, 056
• Chelodina oblonga, 077
Leandro Alcalde
• Hydromedusa tectifera, 113
Abdulaziz R. Alqahtani
• Pelomedusa barbata, 118
Ben Anders
• Mauremys nigricans, 068
Ana Andreu
• Testudo graeca (Western Subspecies Clade), 117
Harald Artner
• Acanthochelys macrocephala, 040
Mehmet K. Atatür
• Rafetus euphraticus, 098
Mark Auliya
• Amyda cartilaginea, 092
Patrick J. Baker
• Cyclanorbis elegans, 089
Dhruvajyoti Basu
• Nilssonia hurum, 048
Jihène Ben Hassine
• Testudo graeca (Western Subspecies Clade), 117
John Sebit Benansio
• Centrochelys sulcata, 110
Rafael Bernhard
• Podocnemis erythrocephala, 087
James F. Berry
• Kinosternon scorpioides, 063
Kristin H. Berry
• Gopherus agassizii, 109
Albert Bertolero
• Testudo hermanni, 059
• Mauremys leprosa, 102
S. Bhupathy
• Hardella thurjii, 023
• Melanochelys trijuga, 038
• Geoclemys hamiltonii, 043
• Indotestudo travancorica, 054
• Lissemys punctata, 076
Emmett L. Blankenship
• Graptemys nigrinoda, 005
Brian C. Bock
• Podocnemis lewyana, 024
• Trachemys callirostris, 042
Stephan Böhm
• Mesoclemmys vanderhaegei, 083
Roger Bour
• Pelusios adansonii, 017
• Pelusios seychellensis, 018
• Turtle Checklist, 000.v4
• Turtle Checklist, 000.v5
• Turtle Checklist, 000.v7
• Fossil Checklist, 000e.v1
• Pelusios castaneus, 095
• Turtle Checklist and Atlas, 000.v8
• Pelusios niger, 105
• Turtle Checklist and Atlas, 000.v9
Jason R. Bourque
• Fossil Checklist, 000e.v1
Ortwin Bourquin
• Pelomedusa subrufa, 007
Deborah S. Bower
• Chelodina expansa, 071
Richard C. Boycott
• Pelusios rhodesianus, 004
• Pelomedusa subrufa, 007
• Pelusios sinuatus, 036
• Kinixys spekii, 115
Bruce J. Brecke
• Emydoidea blandingii, 015
Elizângela S. Brito
• Mesoclemmys vanderhaegei, 083
Donald G. Broadley
• Pelusios rhodesianus, 004
• Pelusios sinuatus, 036
• Cycloderma frenatum, 055
• Kinixys spekii, 115
Rafe M. Brown
• Siebenrockiella leytensis, 066
William S. Brown
• Terrapene coahuila, 049
Kurt A. Buhlmann
• Deirochelys reticularia, 014
Grégory Bulté
* Graptemys geographica, 104
Bernat Burriel-Carranza
• Pelomedusa barbata, 118
R. Bruce Bury
• Actinemys marmorata, 001
Stephen D. Busack
• Mauremys leprosa, 102
Salem M. Busais
• Pelomedusa barbata, 118
James R. Buskirk
• Terrapene nelsoni, 060
Brian P. Butterfield
• Graptemys nigrinoda, 005
Mario R. Cabrera
• Acanthochelys pallidipectoris, 065
• Mesoclemmys vanderhaegei, 083
John Cann
• Myuchelys latisternum, 073
• Elseya lavarackorum, 082
• Myuchelys georgesi, 091
Gladys Cárdenas-Arevalo
• Mesoclemmys dahli, 069
John L. Carr
• Rhinoclemmys nasuta, 034
• Kinosternon dunni, 067
Olga V. Castaño-Mora
• Kinosternon scorpioides albogulare, 064
• Kinosternon dunni, 067
• Mesoclemmys dahli, 069
Adrián Cerdá-Ardura
• Apalone spinifera atra, 021
Eng Heng Chan
• Batagur affinis, 090
Pelf Nyok Chen
• Batagur affinis, 090
Tien-Hsi Chen
• Cuora flavomarginata, 035
Bruce Chessman
• Myuchelys bellii, 088
Marc Cheylan
• Testudo hermanni, 059
Laurent Chirio
• Pelusios castaneus, 095
Justin D. Congdon
• Emydoidea blandingii, 015
• Kinosternon sonoriense, 119
Ben Corey
• Chelodina oblonga, 077
Stefania D’Angelo
• Emys trinacris, 112
Igor G. Danilov
• Fossil Checklist, 000e.v1
Indraneil Das
• Pelochelys cantorii, 011
• Hardella thurjii, 023
• Melanochelys tricarinata, 025
• Chitra indica, 027
• Melanochelys trijuga, 038
• Geoclemys hamiltonii, 043
• Morenia ocellata, 044
• Morenia petersi, 045
• Pangshura sylhetensis, 046
• Nilssonia hurum, 048
• Cuora amboinensis, 053
• Nilssonia leithii, 075
• Cuora mouhotii, 099
Jeffrey E. Dawson
• Mauremys annamensis, 081
• Indotestudo elongata, 096
• Malayemys macrocephala, 108
• Malayemys subtrijuga, 111
Juan M. Daza
• Trachemys callirostris, 042
Neil D’Cruze
• Geochelone elegans, 106
V. Deepak
• Indotestudo travancorica, 054
• Vijayachelys silvatica, 078
Marcelo S. de la Fuente
• Fossil Checklist, 000e.v1
Massimo Delfino
• Fossil Checklist, 000e.v1
Gift Simon Demaya
• Centrochelys sulcata, 110
Anslem de Silva
• Geochelone elegans, 106
Lamine Diagne
• Cyclanorbis senegalensis, 114
Tomas Diagne
• Kinixys homeana, 070
• Kinixys erosa, 084
• Cyclanorbis elegans, 089
• Centrochelys sulcata, 110
• Cyclanorbis senegalensis, 114
Carmen Díaz-Paniagua
• Testudo graeca (Western Subspecies Clade), 117
Arvin C. Diesmos
• Siebenrockiella leytensis, 066
Mae Lowe L. Diesmos
• Siebenrockiella leytensis, 066
James L. Dobie
• Pseudemys alabamensis, 019
C. Kenneth Dodd, Jr.
• Sternotherus depressus, 013
• Kinosternon baurii, 116
J. Sean Doody
• Carettochelys insculpta, 009
Carla Eisemberg
• Carettochelys insculpta, 009
Johannes Els
• Pelomedusa barbata, 118
David Emmett
• Geochelone platynota, 057
Edem A. Eniang
• Centrochelys sulcata, 110
Joshua R. Ennen
• Graptemys pearlensis, 094
Daniel Escoriza
• Testudo graeca (Western Subspecies Clade), 117
• Testudo graeca (Eastern Subspecies Clades), 120
Stephan Ettmar
• Malayemys macrocephala, 108
Michael A. Ewert
• Mauremys annamensis, 081
Renato G. Faria
• Mesoclemmys tuberculata, 097
Phillipa Featherston
• Chelodina burrungandjii, 056
Camila R. Ferrara
• Podocnemis erythrocephala, 087
Darren Fielder
• Myuchelys bellii, 088
Nancy N. FitzSimmons
• Chelodina burrungandjii, 056
Peter S. Floyd
• Pseudemys alabamensis, 019
Damien A. Fordham
• Chelodina oblonga, 077
Germán Forero-Medina
• Kinosternon scorpioides albogulare, 064
• Kinosternon dunni, 067
• Mesoclemmys dahli, 069
Alastair Freeman
• Myuchelys latisternum, 073
• Elseya lavarackorum, 082
Uwe Fritz
• Turtle Checklist and Atlas, 000.v8
• Emys trinacris, 112
• Turtle Checklist and Atlas, 000.v9
• Pelomedusa barbata, 118
Jinzhong Fu
• Cuora flavomarginata, 035
Carlos A. Galvis-Rizo
• Kinosternon dunni, 067
Georgios L. Georgalis
• Fossil Checklist, 000e.v1
Arthur Georges
• Carettochelys insculpta, 009
• Chelodina longicollis, 031
• Chelodina burrungandjii, 056
• Chelodina oblonga, 077
• Myuchelys bellii, 088
• Myuchelys georgesi, 091
• Turtle Checklist and Atlas, 000.v8
• Turtle Checklist and Atlas, 000.v9
Justin Gerlach
• Pelusios castanoides intergularis, 010
• Pelusios subniger parietalis, 016
• Pelusios seychellensis, 018
• Aldabrachelys arnoldi, 028
• Aldabrachelys hololissa, 061
David J. Germano
• Actinemys marmorata, 001
Hanyeh Ghaffari
• Rafetus euphraticus, 098
J. Whitfield Gibbons
• Deirochelys reticularia, 014
• Kinosternon subrubrum, 101
Alan Giraldo
• Rhinoclemmys nasuta, 034
James C. Godwin
• Graptemys nigrinoda, 005
• Graptemys ernsti, 051
• Graptemys pulchra, 072
Gracia Gonzalez-Porter
• Dermatemys mawii, 058
J. Michael Goode
• Mauremys annamensis, 081
Terry E. Graham
• Emydoidea blandingii, 015
Cris Hagen
• Leucocephalon yuwonoi, 039
Adrian Hailey
• Testudo hermanni, 059
• Kinixys spekii, 115
Timo Hartmann
• Indotestudo elongata, 096
Ronnie E. Hatcher
• Mauremys annamensis, 081
Douglas B. Hendrie
• Mauremys annamensis, 081
Emmanuel M. Hema
• Centrochelys sulcata, 110
Tom B. Herman
• Emydoidea blandingii, 015
Ha V. Hoang
• Cuora mouhotii, 099
Kate M. Hodges
• Chelodina longicollis, 031
• Chelodina expansa, 071
Margaretha D. Hofmeyr
• Chersina angulata, 030
Brian D. Horne
• Batagur affinis, 090
Jennifer G. Howeth
• Terrapene coahuila, 049
Daniel F. Hughes
• Kinosternon subrubrum, 101
Flora Ihlow
• Indotestudo elongata, 096
• Malayemys macrocephala, 108
• Malayemys subtrijuga, 111
Bonggi R. Ibarrondo
• Chelodina mccordi, 008
Charles J. Innis
• Leucocephalon yuwonoi, 039
John B. Iverson
• Turtle Checklist, 000.v2
• Turtle Checklist, 000.v3
• Turtle Checklist, 000.v4
• Kinosternon acutum, 062
• Kinosternon scorpioides, 063
• Kinosternon dunni, 067
• Mauremys nigricans, 068
• Turtle Checklist, 000.v5
• Turtle Checklist, 000.v7
• Mauremys annamensis, 081
• Fossil Checklist, 000e.v1
• Turtle Checklist and Atlas, 000.v8
• Kinosternon subrubrum, 101
* Graptemys geographica, 104
• Turtle Checklist and Atlas, 000.v9
• Kinosternon sonoriense, 119
Dale R. Jackson
• Pseudemys concinna, 006
• Deirochelys reticularia, 014
• Pseudemys nelsoni, 041
Hossein Javanbakht
• Testudo graeca (Eastern Subspecies Clades), 120
Gerald R. Johnston
• Kinosternon baurii, 116
Robert L. Jones
• Graptemys oculifera, 033
• Graptemys flavimaculata, 052
• Graptemys pearlensis, 094
Hans-Volker Karl
• Fossil Checklist, 000e.v1
Rod Kennett
• Chelodina longicollis, 031
• Chelodina burrungandjii, 056
• Chelodina oblonga, 077
Khin Myo Myo
• Geochelone platynota, 057
A. Ross Kiester
• Terrapene carolina, 085
Flavius C. Killebrew
• Graptemys versa, 093
Michael W. Klemens
• Kinosternon subrubrum, 101
Yurii V. Kornilev
• Testudo graeca (Eastern Subspecies Clades), 120
Gerald Kuchling
• Chelodina mccordi, 008
Jeffrey W. Lang
• Emydoidea blandingii, 015
Christopher J. Leary
• Pseudemys alabamensis, 019
Peter V. Lindeman
• Sternotherus carinatus, 012
• Graptemys versa, 093
Barbara Livoreil
• Testudo hermanni, 059
Jeffrey E. Lovich
• Graptemys gibbonsi, 029
• Mauremys reevesii, 050
• Graptemys ernsti, 051
• Graptemys pulchra, 072
• Graptemys pearlensis, 094
Luca Luiselli
• Kinixys homeana, 070
• Kinixys erosa, 084
• Cyclanorbis elegans, 089
• Pelusios castaneus, 095
• Pelusios niger, 105
• Centrochelys sulcata, 110
• Cyclanorbis senegalensis, 114
David W. Macdonald
• Geochelone elegans, 106
William E. Magnusson
• Rhinemys rufipes, 079
Patrick Kinyatta Malonza
• Malacochersus tornieri, 107
Thomas M. Mann
• Pseudemys alabamensis, 019
Georgia Mantziou
• Mauremys rivulata, 080
Pablo Marano
• Acanthochelys pallidipectoris, 065
Federico Marrone
• Emys trinacris, 112
Thiago S. Marques
• Mesoclemmys vanderhaegei, 083
• Mesoclemmys tuberculata, 097
Fernando I. Martins
• Hydromedusa maximiliani, 026
Liudmila Mazanaeva
• Testudo graeca (Eastern Subspecies Clades), 120
Timothy E.M. McCormack
• Mauremys annamensis, 081
• Cuora mouhotii, 099
C.J. McCoy
• Graptemys gibbonsi, 029
• Graptemys ernsti, 051
• Graptemys pulchra, 072
Suzanne E. McGaugh
• Apalone spinifera atra, 021
Pearson McGovern
• Cyclanorbis senegalensis, 114
Guido Fabián Medina-Rangel
• Mesoclemmys dahli, 069
Shai Meiri
• Testudo graeca (Eastern Subspecies Clades), 120
Walter E. Meshaka, Jr.
• Kinosternon subrubrum, 101
Daniel O. Mesquita
• Mesoclemmys tuberculata, 097
Sébastien Métrailler
• Acanthochelys macrocephala, 040
• Acanthochelys pallidipectoris, 065
Peter A. Meylan
• Amyda cartilaginea, 092
• Rafetus euphraticus, 098
• Cyclanorbis senegalensis, 114
Russell A. Mittermeier
• Acanthochelys macrocephala, 040
• Podocnemis erythrocephala, 087
Don Moll
• Dermatemys mawii, 058
• Malacochersus tornieri, 107
Edward O. Moll
• Batagur baska, 037
• Batagur affinis, 090
• Amyda cartilaginea, 092
Anirhuddha Mookerjee
• Geochelone elegans, 106
Geraldo J.B. Moura
• Mesoclemmys tuberculata, 097
Robert W. Murphy
• Gopherus agassizii, 109
B.H.C.K. Murthy
• Nilssonia leithii, 075
Reginald T. Mwaya
• Malacochersus tornieri, 107
David H. Nelson
• Pseudemys alabamensis, 019
Jacob Mueti Ngwava
• Malacochersus tornieri, 107
Fabrizio Oneto
• Emys trinacris, 112
Hidetoshi Ota
• Geoemyda japonica, 002
• Mauremys japonica, 003
• Cuora flavomarginata, 035
• Geoemyda spengleri, 047
• Mauremys reevesii, 050
Dario Ottonello
• Emys trinacris, 112
Vivian P. Páez
• Podocnemis lewyana, 024
• Trachemys callirostris, 042
Charles W. Painter
• Pseudemys gorzugi, 100
Michael J. Pappas
• Emydoidea blandingii, 015
James F. Parham
• Turtle Checklist, 000.v1
• Turtle Checklist, 000.v2
Lorena Paszko
• Acanthochelys pallidipectoris, 065
Fabio Petrozzi
• Pelusios castaneus, 095
• Pelusios niger, 105
• Centrochelys sulcata, 110
Leland J.S. Pierce
• Pseudemys gorzugi, 100
Kalyar Platt
• Batagur baska, 037
• Geochelone platynota, 057
• Chitra vandijki, 074
• Batagur affinis, 090
Steven G. Platt
• Rhinoclemmys areolata, 022
• Batagur baska, 037
• Leucocephalon yuwonoi, 039
• Geochelone platynota, 057
• Chitra vandijki, 074
• Batagur affinis, 090
• Malayemys subtrijuga, 111
John R. Polisar
• Dermatemys mawii, 058
Paulino Ponce-Campos
• Terrapene nelsoni, 060
Georgi Popgeorgiev
• Testudo graeca (Eastern Subspecies Clades), 120
Peter Praschag
• Batagur baska, 037
• Pangshura sylhetensis, 046
• Lissemys punctata, 076
• Vijayachelys silvatica, 078
• Batagur affinis, 090
Peter C.H. Pritchard
• Chelus fimbriata, 020
• Hydromedusa tectifera, 113
Thomas R. Rainwater
• Rhinoclemmys areolata, 022
• Geochelone platynota, 057
• Chitra vandijki, 074
Madhuri Ramesh
• Indotestudo travancorica, 054
Luis E. Rentería-Moreno
• Kinosternon dunni, 067
Adriana Restrepo
• Podocnemis lewyana, 024
Anders G.J. Rhodin
• Turtle Checklist, 000.v1
• Chelodina mccordi, 008
• Acanthochelys macrocephala, 040
• Turtle Checklist, 000.v2
• Turtle Checklist, 000.v3
• Turtle Checklist, 000.v4
• Turtle Checklist, 000.v5
• Turtle Checklist, 000.v7
• Fossil Checklist, 000e.v1
• Turtle Checklist and Atlas, 000.v8
• Turtle Checklist and Atlas, 000.v9
Enrique Richard
• Acanthochelys pallidipectoris, 065
Lina Rifai
• Mauremys rivulata, 080
John Roe
• Chelodina longicollis, 031
Mark Rose
• Carettochelys insculpta, 009
Philip C. Rosen
• Kinosternon sonoriense, 119
Walter Sachsse
• Cycloderma frenatum, 055
Rocio Maria Sánchez
• Hydromedusa tectifera, 113
Daniel O. Santana
• Mesoclemmys tuberculata, 097
Chuck Schaffer
• Manouria emys, 086
Rick Schaffer
• Manouria emys, 086
Sabine Schoppe
• Cuora amboinensis, 053
Gabriel H. Segniagbeto
• Pelusios castaneus, 095
• Pelusios niger, 105
• Centrochelys sulcata, 110
Will Selman
• Graptemys gibbonsi, 029
• Graptemys oculifera, 033
• Graptemys flavimaculata, 052
Saibal Sengupta
• Morenia petersi, 045
• Pangshura sylhetensis, 046
H. Bradley Shaffer
• Turtle Checklist, 000.v3
• Turtle Checklist, 000.v4
• Turtle Checklist, 000.v5
• Turtle Checklist, 000.v7
• Fossil Checklist, 000e.v1
• Turtle Checklist and Atlas, 000.v8
• Turtle Checklist and Atlas, 000.v9
Uri Shanas
• Testudo graeca (Eastern Subspecies Clades), 120
Mohammed Shobrak
• Pelomedusa barbata, 118
Shailendra Singh
• Chitra indica, 027
• Nilssonia hurum, 048
Shashwat Sirsi
• Nilssonia leithii, 075
Francisco Soberón-Mobarak
• Apalone spinifera atra, 021
Sitha Som
• Indotestudo elongata, 096
Franco L. Souza
• Hydromedusa maximiliani, 026
Filippo Spadola
• Emys trinacris, 112
Stephen Spawls
• Kinixys spekii, 115
Ricky-J. Spencer
• Myuchelys georgesi, 091
Craig B. Stanford
• Manouria emys, 086
Marie E.B. Stone
• Kinosternon sonoriense, 119
Paul A. Stone
• Kinosternon sonoriense, 119
Richard P.J.H. Struijk
• Cuora mouhotii, 099
James N. Stuart
• Trachemys gaigeae, 032
• Graptemys versa, 093
• Pseudemys gorzugi, 100
• Kinosternon sonoriense, 119
Thanda Swe
• Geochelone platynota, 057
Emerson Y. Sy
• Siebenrockiella leytensis, 066
Akio Takahashi
• Fossil Checklist, 000e.v1
Ertan Taskavak
• Rafetus euphraticus, 098
Kumthorn Thirakhupt
• Manouria emys, 086
• Malayemys macrocephala, 108
Scott Thomson
• Chelodina burrungandjii, 056
• Elseya lavarackorum, 082
• Fossil Checklist, 000e.v1
Anton Tucker
• Chelodina burrungandjii, 056
Oguz Türkozan
• Testudo graeca (Eastern Subspecies Clades), 120
Elias Tzoras
• Testudo graeca (Eastern Subspecies Clades), 120
Peter Paul van Dijk
• Turtle Checklist, 000.v1
• Batagur baska, 037
• Turtle Checklist, 000.v2
• Turtle Checklist, 000.v3
• Turtle Checklist, 000.v4
• Turtle Checklist, 000.v5
• Turtle Checklist, 000.v7
• Fossil Checklist, 000e.v1
• Batagur affinis, 090
• Amyda cartilaginea, 092
• Cuora mouhotii, 099
• Turtle Checklist and Atlas, 000.v8
• Malayemys macrocephala, 108
• Turtle Checklist and Atlas, 000.v9
Mario Vargas-Ramirez
• Podocnemis lewyana, 024
Karthikeyan Vasudevan
• Indotestudo travancorica, 054
• Nilssonia leithii, 075
• Vijayachelys silvatica, 078
Luciano M. Verdade
• Mesoclemmys vanderhaegei, 083
Gustavo H.C. Vieira
• Mesoclemmys tuberculata, 097
Sabine Vinke
• Acanthochelys macrocephala, 040
• Acanthochelys pallidipectoris, 065
Thomas Vinke
• Acanthochelys macrocephala, 040
• Acanthochelys pallidipectoris, 065
Richard C. Vogt
• Apalone spinifera atra, 021
• Rhinoclemmys areolata, 022
• Dermatemys mawii, 058
• Kinosternon acutum, 062
• Rhinemys rufipes, 079
• Podocnemis erythrocephala, 087
• Graptemys ouachitensis, 103
* Graptemys geographica, 104
Raju Vyas
• Geochelone elegans, 106
Pratyaporn Wanchai
• Manouria emys, 086
Joseph P. Ward
• Pseudemys concinna, 006
• Trachemys gaigeae, 032
• Pseudemys gorzugi, 100
Robert G. Webb
• Lissemys punctata, 076
Michael Welsh
• Myuchelys georgesi, 091
Ronald E. Willemsen
• Testudo hermanni, 059
Lisabeth L. Willey
• Terrapene carolina, 085
Dawn S. Wilson
• Kinosternon baurii, 116
Win Ko Ko
• Geochelone platynota, 057
• Chitra vandijki, 074
Takashi Yabe
• Mauremys japonica, 003
Yuichirou Yasukawa
• Geoemyda japonica, 002
• Mauremys japonica, 003
• Cuora flavomarginata, 035
• Geoemyda spengleri, 047
• Mauremys reevesii, 050
Marco Alberto Luca Zuffi
• Emys trinacris, 112