Contributing Photographers
Contributing CBFTT Account Photographers
(not including TTWG Checklist and Atlas photographers)
We encourage any and all who have good photos of freshwater turtles and tortoises to consider submitting them to us for consideration of inclusion in upcoming new accounts or eventually revised older accounts. See specific instructions for Contributing Photographers. All photos used will be acknowledged in the published accounts and here on this page. Thank you.
To date, the following individuals have contributed photographs for published accounts in this on-going project.
Number of contributing photographers published to date: 293
Number of accounts published to date: 126
* = Also an author or co-author of at least one account; see author list
Collette Adams
• Terrapene carolina, 08
Godfrey C. Akani *
• Pelusios niger, 105
Fatehi Al Saaw
• Pelomedusa barbata, 118
Leandro Alcalde *
• Hydromedusa tectifera, 113
Patrick Alexander
• Terrapene ornata, 126
Abdulaziz R. Alqahtani *
• Pelomedusa barbata, 118
Bhaba Amatya
• Melanochelys trijuga, 038
Ben Anders *
• Mauremys nigricans, 068
Mehmet K. Atatür *
• Rafetus euphraticus, 098
Markus Auer
• Mauremys caspica, 125
Mark Auliya *
• Manouria emys, 086
• Amyda cartilaginea, 092
Tracy Baker
• Cyclanorbis elegans, 089
Richard D. Bartlett
• Sternotherus carinatus, 012
• Pseudemys nelsoni, 041
• Kinosternon subrubrum, 101
Chittaranjan Baruah
• Chitra indica, 027
• Geoclemys hamiltonii, 043
• Nilssonia hurum, 048
• Lissemys punctata, 076
Jeff Beane
• Kinosternon baurii, 116
Virginia C.D. Bernardes
• Podocnemis erythrocephala, 087
Rafael Bernhard *
• Podocnemis erythrocephala, 087
Kristin H. Berry *
• Gopherus agassizii, 109
Albert Bertolero *
• Testudo hermanni, 059
• Mauremys leprosa, 102
S. Bhupathy *
• Melanochelys trijuga, 038
Michael S. Bogle
• Terrapene nelsoni, 060
Roger Bour *
• Pelusios adansonii, 017
• Pelusios seychellensis, 018
• Pelusios castaneus, 095
Deborah S. Bower *
• Chelodina expansa, 071
Richard C. Boycott *
• Pelusios rhodesianus, 004
• Pelusios sinuatus, 036
• Kinixys spekii, 115
Greg Brashear
• Kinosternon baurii, 116
Bruce Brecke
• Graptemys ouachitensis, 103
Raissa Bressan
• Hydromedusa tectifera, 113
Henrik Bringsøe *
• Testudo marginata, 122
Elizângela S. Brito *
• Mesoclemmys vanderhaegei, 083
Rafe M. Brown *
• Siebenrockiella leytensis, 066
William S. Brown *
• Terrapene coahuila, 049
Kurt A. Buhlmann *
• Deirochelys reticularia, 014
• Kinosternon baurii, 116
Grégory Bulté *
• Graptemys geographica, 104
Betty L. Burge
• Gopherus agassizii, 109
R. Bruce Bury *
• Actinemys marmorata, 001
Stephen D. Busack *
• Mauremys leprosa, 102
James R. Buskirk *
• Rhinoclemmys areolata, 022
Brian O. Butler
• Emydoidea blandingii, 015
Howard G. Byrne
• Kinosternon sonoriense, 119
Alejandra Cadavid
• Podocnemis lewyana, 024
Matt Cage
• Terrapene nelsoni, 060
Young Cage
• Terrapene nelsoni, 060
John Cann *
• Carettochelys insculpta, 009
• Chelodina longicollis, 031
• Myuchelys latisternum, 073
• Chelodina oblonga, 077
• Elseya lavarackorum, 082
• Myuchelys georgesi, 091
Gladys Cárdenas-Arevalo *
• Mesoclemmys dahli, 069
John L. Carr *
• Rhinoclemmys nasuta, 034
• Rhinoclemmys annulata, 123
Luis Ceriaco
• Pelusios castaneus, 095
Jan Červenka
• Pelomedusa barbata, 118
Eng Heng Chan *
• Batagur affinis, 090
Tien-Hsi Chen *
• Cuora flavomarginata, 035
Laurent Chirio
• Kinixys erosa, 084
Ana Colima
• Terrapene ornata, 126
Suzanne Collins
• Terrapene ornata, 126
Justin D. Congdon *
• Emydoidea blandingii, 015
• Kinosternon sonoriense, 119
Arturo Conteras-Arquieta
• Terrapene coahuila, 049
Colin Croft
• Terrapene ornata, 126
Stefania D’Angelo *
• Emys trinacris, 112
Indraneil Das *
• Pelochelys cantorii, 011
• Hardella thurjii, 023
• Melanochelys tricarinata, 025
• Melanochelys trijuga, 038
• Geoclemys hamiltonii, 043
• Morenia ocellata, 044
• Morenia petersi, 045
• Pangshura sylhetensis, 046
• Nilssonia hurum, 048
• Cuora amboinensis, 053
• Lissemys punctata, 076
• Cuora mouhotii, 099
Shekar Dattatri
• Chitra indica, 027
Jeffrey E. Dawson *
• Mauremys annamensis, 081
• Indotestudo elongata, 096
• Malayemys subtrijuga, 111
• Terrapene ornata, 126
Neil D’Cruze *
• Geochelone elegans, 106
V. Deepak *
• Indotestudo travancorica, 054
• Nilssonia leithii, 075
• Vijayachelys silvatica, 078
Pablo R. Delis
• Kinosternon subrubrum, 101
Anslem de Silva *
• Geochelone elegans, 106
Desert Tortoise Preserve Committee
• Gopherus agassizii, 109
Bernard Devaux
• Terrapene nelsoni, 060
Tomas Diagne *
• Kinixys homeana, 070
• Kinixys erosa, 084
• Pelusios niger, 105
• Centrochelys sulcata, 110
Carmen Díaz-Paniagua *
• Testudo graeca (Western Subspecies Clade), 117
James L. Dobie *
• Pseudemys alabamensis, 019
C. Kenneth Dodd, Jr. *
• Sternotherus depressus, 013
• Kinosternon baurii, 116
Jonathan Duran
• Kinosternon sonoriense, 119
Carla Eisemberg *
• Carettochelys insculpta, 009
Johannes Els *
• Pelomedusa barbata, 118
• Mauremys caspica, 125
David Emmett *
• Amyda cartilaginea, 092
Joshua R. Ennen *
• Graptemys pearlensis, 094
Daniel Escoriza *
• Testudo graeca (Western Subspecies Clade), 117
• Testudo marginata, 122
Michael A. Ewert *
• Deirochelys reticularia, 014
Alejandro Fallabrino
• Hydromedusa tectifera, 113
Shirley Famelli
• Hydromedusa maximiliani, 026
Darren Fielder *
• Myuchelys bellii, 088
• Myuchelys georgesi, 091
Paulo L. Filho
• Hydromedusa maximiliani, 026
Nancy N. FitzSimmons *
• Chelodina burrungandjii, 056
Florida Fish and Wildlife Commission
• Kinosternon baurii, 116
Karl Ford
• Kinosternon sonoriense, 119
Damien A. Fordham *
• Chelodina oblonga, 077
Germán Forero-Medina *
• Kinosternon scorpioides albogulare, 064
• Kinosternon dunni, 067
• Mesoclemmys dahli, 069
Michael R.J. Forstner
• Pseudemys gorzugi, 100
Alastair Freeman *
• Myuchelys latisternum, 073
• Elseya lavarackorum, 082
Peter Geissler
• Indotestudo elongata, 096
Arthur Georges *
• Carettochelys insculpta, 009
• Chelodina longicollis, 031
• Chelodina burrungandjii, 056
• Chelodina oblonga, 077
• Myuchelys bellii, 088
• Myuchelys georgesi, 091
Justin Gerlach *
• Pelusios castanoides intergularis, 010
• Pelusios subniger parietalis, 016
• Aldabrachelys arnoldi, 028
• Aldabrachelys hololissa, 061
David J. Germano *
• Actinemys marmorata, 001
• Terrapene ornata, 126
Hanyeh Ghaffari *
• Rafetus euphraticus, 098
Tom Gill
• Terrapene ornata, 126
Alan Giraldo *
• Rhinoclemmys nasuta, 034
James C. Godwin *
• Graptemys nigrinoda, 005
• Graptemys ernsti, 051
• Graptemys pulchra, 072
Egbert Golz
• Testudo marginata, 122
Gracia Gonzalez-Porter *
• Dermatemys mawii, 058
Terry E. Graham *
• Emydoidea blandingii, 015
Sally Grundy
• Chelodina longicollis, 031
Ferry Grünewald
• Pelusios niger, 105
• Rhinoclemmys annulata, 123
Wulf Haacke
• Cycloderma frenatum, 055
Cris Hagen *
• Melanochelys trijuga, 038
• Leucocephalon yuwonoi, 039
Adrian Hailey *
• Testudo hermanni, 059
William M. Hammond
• Kinosternon sonoriense, 119
James H. Harding
• Sternotherus carinatus, 012
• Emydoidea blandingii, 015
• Rhinoclemmys areolata, 022
• Graptemys pseudogeographica, 124
Jiri Haureljuk
• Kinixys spekii, 115
George Heinrich
• Deirochelys reticularia, 014
Dean A. Hendrickson
• Terrapene coahuila, 049
Mario Herz
• Mauremys caspica, 125
Terry Hibbitts
• Graptemys versa, 093
Pierson Hill
• Pseudemys nelsoni, 041
Lina Hinestroza
• Podocnemis lewyana, 024
Kate M. Hodges *
• Chelodina expansa, 071
Margaretha D. Hofmeyr *
• Chersina angulata, 030
Dan C. Holland
• Actinemys marmorata, 001
Brian D. Horne
• Terrapene ornata, 126
Janet Hostetter
• Emydoidea blandingii, 015
Jennifer G. Howeth *
• Terrapene coahuila, 049
Rick Hudson
• Batagur trivittata, Cover Photo
• Batagur baska, 037
Flora Ihlow *
• Indotestudo elongata, 096
• Malayemys macrocephala, 108
• Malayemys subtrijuga, 111
• Kinixys spekii, 115
Iriomote Wildlife Conservation Center
• Cuora flavomarginata, 035
Steve Ishii
• Gopherus agassizii, 109
John B. Iverson *
• Rhinoclemmys areolata, 022
• Trachemys gaigeae, 032
• Terrapene nelsoni, 060
• Kinosternon acutum, 062
• Kinosternon scorpioides, 063
• Mauremys nigricans, 068
• Kinosternon subrubrum, 101
• Graptemys geographica, 104
• Kinosternon baurii, 116
• Kinosternon sonoriense, 119
• Kinosternon flavescens, 121
• Graptemys pseudogeographica, 124
Dale R. Jackson *
• Pseudemys concinna, 006
Hossein Javanbakht *
• Testudo graeca (Eastern Subspecies Clades), 120
S. Jayakumar
• Melanochelys trijuga, 038
Karen A. Jensen
• Amyda cartilaginea, 092
Jason M. Jones
• Terrapene coahuila, 049
Michael T. Jones
• Terrapene carolina, 085
Robert L. Jones *
• Graptemys oculifera, 033
• Graptemys pearlensis, 094
Nobuhiro Kawazoe
• Geoemyda spengleri, 047
• Mauremys reevesii, 050
C.K. Kelly
• Terrapene ornata, 126
Anser Khan
• Lissemys punctata, 076
Heng Kimchhay
• Batagur affinis, 090
John Kleopfer
• Kinosternon baurii, 116
Gunther Köhler
• Mesoclemmys vanderhaegei, 083
Lukáš Kratochvíl
• Pelomedusa barbata, 118
Gerald Kuchling *
• Melanochelys trijuga, 038
• Morenia ocellata, 044
• Chitra vandijki, 074
• Amyda cartilaginea, 092
Brian Lang
• Pseudemys gorzugi, 100
Thomas Leuteritz
• Kinixys erosa, 084
Peter V. Lindeman *
• Graptemys flavimaculata, 052
• Graptemys versa, 093
• Graptemys ouachitensis, 103
• Graptemys pseudogeographica, 124
Scott Loarie
• Terrapene ornata, 126
Victor Loehr
• Kinixys spekii, 115
Paula C. Lopes
• Mesoclemmys vanderhaegei, 083
Nora López-León
• Kinosternon scorpioides, 063
Bill Love
• Graptemys gibbonsi, 029
Jeffrey E. Lovich *
• Graptemys gibbonsi, 029
Gary Luciano
• Pseudemys nelsoni, 041
Luca Luiselli *
• Kinixys homeana, 070
• Kinixys erosa, 084
• Pelusios castaneus, 095
• Pelusios niger, 105
• Centrochelys sulcata, 110
William E. Magnusson *
• Rhinemys rufipes, 079
Thomas M. Mann *
• Pseudemys alabamensis, 019
Georgia Mantziou *
• Mauremys rivulata, 080
Maurizio Marchese
• Emys trinacris, 112
Federico Marrone *
• Emys trinacris, 112
Thiago S. Marques *
• Mesoclemmys vanderhaegei, 083
Ramon Mascort
• Terrapene coahuila, 049
Mark Massar
• Gopherus agassizii, 109
Brooke N. Massengill
• Kinosternon sonoriense, 119
Jeremy K. Massengill
• Kinosternon sonoriense, 119
Carl D. May
• Kinosternon baurii, 116
Liudmila F. Mazanaeva *
• Testudo graeca (Eastern Subspecies Clades), 120
• Mauremys caspica, 125
Timothy McCormack *
• Mauremys annamensis, 081
• Cuora mouhotii, 099
Suzanne E. McGaugh *
• Apalone spinifera atra, 021
Pearson McGovern
• Cyclanorbis senegalensis, 114
Ann McLuckie
• Gopherus agassizii, 109
Konrad Mebert
• Kinixys erosa, 084
Pratheep Meewattana
• Malayemys macrocephala, 108
Fernanda Melo
• Acanthochelys macrocephala, 040
Khaled Merabet
• Mauremys leprosa, 102
Melvin Mérida
• Dermatemys mawii, 058
Sébastien Métrailler *
• Acanthochelys macrocephala, 040
• Acanthochelys pallidipectoris, 065
Peter A. Meylan
• Pseudemys concinna, 006
• Cyclanorbis senegalensis, 114
Marsha Miller
• Terrapene coahuila, 049
Russell A. Mittermeier *
• Chelus fimbriata, 020
• Acanthochelys macrocephala, 040
• Dermatemys mawii, 058
• Podocnemis erythrocephala, 087
Asghar Mobaraki
• Mauremys caspica, 125
Mojave Natural Preserve
• Gopherus agassizii, 109
Don Moll *
• Malacochersus tornieri, 107
Edward O. Moll *
• Batagur affinis, 090
Robert H. Mount *
• Pseudemys alabamensis, 019
Museo de Herpetología de la Universidad de Antioquia
• Trachemys callirostris, 042
Reginald T. Mwaya *
• Malacochersus tornieri, 107
Rahul Naik
• Nilssonia leithii, 075
Nguyen Thanh Luan
• Cuora mouhotii, 099
Nguyen Van Sang
• Geoemyda spengleri, 047
Andreas Nöllert
• Mauremys leprosa, 102
• Hydromedusa tectifera, 113
Greg Norsworthy
• Pseudemys nelsoni, 041
Andy Noss
• Kinixys erosa, 084
Jennifer Wallin Oliver
• Trachemys gaigeae, 032
Annette Olsson
• Pelochelys cantorii, 011
Dario Ottonello *
• Emys trinacris, 112
Mathieu Ouellette
• Graptemys geographica, 104
Vivian P. Páez *
• Podocnemis lewyana, 024
• Trachemys callirostris, 042
Charles W. Painter *
• Trachemys gaigeae, 032
• Pseudemys gorzugi, 100
James F. Parham *
• Hardella thurjii, 023
Fred Parker
• Chelodina oblonga, 077
Lorena Paszko *
• Acanthochelys pallidipectoris, 065
John Pemberton
• Aldabrachelys arnoldi, 028
Jarmo Perälä
• Cuora flavomarginata, 035
Fabio Petrozzi *
• Pelusios castaneus, 095
• Centrochelys sulcata, 110
Hans-Dieter Philippen
• Cuora flavomarginata, 035
• Melanochelys trijuga, 038
• Morenia petersi, 045
• Cuora amboinensis, 053
Kalyar Platt *
• Cuora amboinensis, 053
• Geochelone platynota, 057
Håkan Pohlstrand
• Centrochelys sulcata, 110
• Cyclanorbis senegalensis, 114
James Poindexter
• Terrapene nelsoni, 060
Paulino Ponce-Campos *
• Terrapene nelsoni, 060
Georgi Popgeorgiev *
• Testudo graeca (Eastern Subspecies Clades), 120
Frank Portillo
• Terrapene ornata, 126
Peter Praschag *
• Batagur baska, 037
• Pangshura sylhetensis, 046
• Lissemys punctata, 076
• Vijayachelys silvatica, 078
• Geochelone elegans, 106
Peter C.H. Pritchard *
• Chelus fimbriata, 020
• Trachemys callirostris, 042
Evan S.H. Quah
• Malayemys macrocephala, 108
S.M.A. Rashid
• Batagur baska, 037
Freya Reder
• Gopherus agassizii, 109
Michael Redmer
• Rhinoclemmys annulata, 123
Paul Reef
• Morenia ocellata, 044
Anders G.J. Rhodin *
• Pelomedusa subrufa, 007
• Chelodina mccordi, 008
• Chelus fimbriata, 020
• Melanochelys trijuga, 038
• Acanthochelys macrocephala, 040
• Kinixys homeana, 070
• Chelodina oblonga, 077
• Mauremys rivulata, 080
• Malacochersus tornieri, 107
• Mauremys caspica, 125
Enrique Richard *
• Acanthochelys pallidipectoris, 065
Casey Richart
• Terrapene ornata, 126
Lina Rifai *
• Mauremys rivulata, 080
Maurice Rodrigues
• Cyclanorbis elegans, 089
John Roe *
• Chelodina longicollis, 031
Cameron Rognan
• Gopherus agassizii, 109
Oliver Römpp
• Chelodina longicollis, 031
José Vicente Rueda-Almonacid
• Rhinoclemmys nasuta, 034
Barbod Safaei-Mahroo
• Rafetus euphraticus, 098
San Diego Zoo Global
• Gopherus agassizii, 109
Suzy Sanoja-Sarabia
• Pseudemys gorzugi, 100
Daniel O. Santana •
• Mesoclemmys tuberculata, 097
Patty Scanlan
• Terrapene coahuila, 049
• Terrapene carolina, 085
Rita Scardino
• Emys trinacris, 112
Chuck Schaffer *
• Manouria emys, 086
Rick Schaffer *
• Manouria emys, 086
Fabian Schmidt
• Malacochersus tornieri, 107
Christoph Schneider
• Testudo graeca (Western Subspecies Clade), 117
Willi Schneider
• Testudo graeca (Western Subspecies Clade), 117
Sabine Schoppe *
• Cuora amboinensis, 053
• Siebenrockiella leytensis, 066
Jake Scott
• Kinosternon baurii, 116
Gabriel Hoinsoudé Segniagbeto *
• Pelusios castaneus, 095
Will Selman *
• Graptemys gibbonsi, 029
• Graptemys flavimaculata, 052
Uri Shanas *
• Testudo graeca (Eastern Subspecies Clades), 120
Liron Shapira
• Testudo graeca (Eastern Subspecies Clades), 120
Rowan Simon
• Myuchelys georgesi, 091
Shailendra Singh *
• Morenia petersi, 045
• Nilssonia hurum, 048
• Lissemys punctata, 076
Shashwat Sirsi *
• Nilssonia leithii, 075
Alex Slavenko
• Testudo graeca (Eastern Subspecies Clades), 120
Joaquim Soler Massana
• Mauremys leprosa, 102
Sitha Som *
• Indotestudo elongata, 096
Franco L. Souza *
• Hydromedusa maximiliani, 026
Stephen Spawls *
• Cycloderma frenatum, 055
• Kinixys spekii, 115
Craig B. Stanford *
• Manouria emys, 086
Brian D. Stanila
• Kinosternon sonoriense, 119
Brett Stearns
• Pseudemys gorzugi, 100
Dirk Stevenson
• Kinosternon baurii, 116
Bev Steveson
• Gopherus agassizii, 109
Kevin Stohlgren
• Kinosternon baurii, 116
Marie E.B. Stone *
• Kinosternon sonoriense, 119
Paul A. Stone *
• Kinosternon sonoriense, 119
Richard P.J.H. Struijk *
• Cuora mouhotii, 099
James N. Stuart *
• Trachemys gaigeae, 032
• Kinosternon sonoriense, 119
Emerson Y. Sy *
• Siebenrockiella leytensis, 066
Steffen Szymanski
• Malayemys subtrijuga, 111
Jack Thompson
• Kinosternon baurii, 116
Robert C. Thomson
• Sternotherus carinatus, 012
Stanley E. Trauth
• Sternotherus carinatus, 012
• Graptemys pseudogeographica, 124
Claire Treilibs
• Chelodina expansa, 071
Apostolis Trichas
• Mauremys rivulata, 080
Ashutosh Tripathi
• Chitra indica, 027
Truong Nguyen Quang
• Geoemyda spengleri, 047
Anton Tucker *
• Chelodina burrungandjii, 056
Michael Tuma
• Gopherus agassizii, 109
Oguz Türkozan *
• Testudo graeca (Eastern Subspecies Clades), 120
Turtle Conservancy
• Malacochersus tornieri, 107
Elias Tzoras *
• Testudo graeca (Eastern Subspecies Clades), 120
• Testudo marginata, 122
U.S. Geological Survey
• Gopherus agassizii, 109
Melita Vamberger
• Emys trinacris, 112
Thomas R. Van Devender
• Pseudemys gorzugi, 100
Peter Paul van Dijk *
• Melanochelys trijuga, 038
• Geoclemys hamiltonii, 043
• Geochelone platynota, 057
• Mauremys annamensis, 081
• Terrapene carolina, 085
• Geochelone elegans, 106
Karthik Vasudevan *
• Nilssonia leithii, 075
• Vijayachelys silvatica, 078
Jay Villemarette
• Cyclanorbis elegans, 089
Thomas and Sabine Vinke *
• Acanthochelys macrocephala, 040
• Acanthochelys pallidipectoris, 065
• Mesoclemmys vanderhaegei, 083
John Visser
• Trachemys callirostris, 042
Evangelos Vlachos
• Testudo marginata, 122
Richard C. Vogt *
• Rhinoclemmys areolata, 022
• Dermatemys mawii, 058
• Kinosternon acutum, 062
• Rhinemys rufipes, 079
• Podocnemis erythrocephala, 087
• Graptemys ouachitensis, 103
Raju Vyas *
• Geochelone elegans, 106
Pratyaporn Wanchai *
• Manouria emys, 086
Joseph P. Ward *
• Pseudemys concinna, 006
• Pseudemys gorzugi, 100
Peter Wellings
• Chelodina oblonga, 077
Ronald E. Willemsen *
• Testudo hermanni, 059
Lisabeth L. Willey *
• Terrapene carolina, 085
Dawn S. Wilson *
• Kinosternon baurii, 116
Win Ko Ko *
• Chitra vandijki, 074
WWF Italia
• Emys trinacris, 112
Yuichirou Yasukawa *
• Geoemyda japonica, 002
• Mauremys japonica, 003
• Cuora flavomarginata, 035
• Geoemyda spengleri, 047
George R. Zug
• Morenia ocellata, 044